Mainly by two methods: Growth “rings” on scales, and/or ringlike structures found in otoliths (small bones of the inner ear), are examined and counted. The rings correspond to seasonal changes in the environment and can be compared to the annual rings of tree trunks. A series of fine rings are laid down in scales for each year of life in summer, the rings grow faster and have relatively wide separations; in winter, slower growth is indicated by narrow separations between rings. Each pair of rings indicates one year. Because scale rings are sometimes influenced by other factors, scientists often use otoliths, whose ringlike structures also indicate years of life.
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